Tribe Brachyglutini Raffray, 1904
- Brachyglutini Raffray, A. 1904. Genera et catalogue des Psélaphides. Annales de la Société Entomologique de France 73: 1-400 [108].
Type genus:
Brachygluta Thomson, 1859. - Bryaxes LeConte, J.L. 1861. Classification of the Coleoptera of North America. Prepared for the Smithsonian Institution. Part I. Smithsonian Miscellaneous Collections 3: xxiv + 286 pp. [Date published 1861-1862] [57] [based on rejected generic name, Bryaxis Leach, 1817].
Introduction
This tribe currently holds 4 subtribes with 116 genera. The distinctive subtribes Baradina Park, Decarthrina Park, and Eupseniina Park are restricted to the New World, and are most diverse in the Neotropics. All other genera are placed in the Brachyglutina, which now includes all members of the former subtribes Pselaptina and Halorabyxina, as well as the Globina.
Twenty-eight genera of Brachyglutini occur in Australia. Four of these genera are also found in New Zealand (Startes, Anabaxis, Eupines, Rybaxis), with the first two found only in New Zealand and Australia. Five genera are also found in the Oriental Region (Eupines, Batraxis, Pedisinops, Physoplectus, Briara), and two genera are cosmopolitan (Rybaxis, Reichenbachia). Nineteen genera are known only from Australia, though this probably reflects to some extent the very poor knowledge of the brachyglutine fauna of the Oriental Region (Chandler 2001: 290).
Diagnosis
Head with antennomeres I short; ocular-mandibular carinae present; with 10-11 antennomeres; third segments of maxillary palpi short and triangular, fourth segments ovoidal to elongate-conical (unless modified in males), usually obliquely truncate at bases. Prothorax with paranotal carinae. Legs with second tarsomeres as long as or longer than third tarsomeres. Abdominal margins angulate, only a few genera with paratergites very narrow or fused to form rounded lateral margins, paratergites usually clearly visible for visible tergites 1-3 (IV-VI); visible sternite 1 (III) narrowly visible at base between metacoxae and laterally, 2 (IV) much longer than first at midline. Aedeagus usually symmetric, with parameres similar in form; some genera with relatively few foveae and sulci have asymmetric aedeagi.
ID Keys
Key to Australian genera of Brachyglutini (Chandler 2001: 292).
1. Elytral flanks with both subhumeral fovea and marginal stria … 2
Elytra lacking subhumeral fovea and marginal stria, rarely with longitudinal sulcus on apical half of elytral flanks … 3
2. (1) Pronotum with angular to curved transverse antebasal sulcus, median antebasal fovea absent to as large as lateral antebasal foveae; ventrolateral margins of head rounded beneath eyes … Rybaxis Saulcy
Pronotum lacking any trace of antebasal sulcus, setose median antebasal fovea as large as lateral antebasal foveae; head tuberculate beneath eyes … Physoplectus Reitter
3. (1) Antennal clubs with antennomeres IX and X slightly elongate, both narrowing to apices in apical halves, X larger than IX but clubs appearing formed solely from antennomeres XI; head with setose pits beneath eyes; pronotum lacking median antebasal fovea, lateral antebasal foveae faint or lacking … Booloumba Chandler
Antennae with antennomeres IX and X usually different in form, if similar (some female Eupines) then lacking setose pits beneath eyes, IX and X usually clearly transverse, quadrate or elongate, not narrowing to apices, clubs usually formed by apical 2-3 segments; setose pits beneath eyes found only in Woodenbong and Wollomombi; presence of pronotal foveae variable … 4
4. (3) Head with dorsal postantennal pits; pronotum often with antebasal sulcus weakly indicated at least near antebasal foveae … 5
Head lacking dorsal postantennal pits, though postantennal slits may extend medially from lateral margins of head (Wiangaree); pronotum with antebasal sulcus lacking (except Paluma) … 6
5. (4) Gula swollen at middle, in lateral view angularly rounded toward gular foveae; pronotum with antebasal sulcus often faint or feebly indicated near median antebasal fovea, this fovea distinct … Anabaxis Raffray
Gula gently rounded in lateral view to nearly flat; pronotum with faint to strong antebasal sulcus, median antebasal fovea often no wider than sulcus and indistinct … Mallanganee Chandler
6. (4) Head lacking vertexal foveae, vertexal disc simple … 7
Head with vertexal foveae present, if vertex obviously modified (males) then foveae may be obscured … 9
7. (6) All antennomeres elongate, slightly laterally flattened, antennal clubs indistinct; metasternal length between meso- and metacoxae subequal to mesocoxal length … Araneabaxis Chandler
Intermediate antennomeres clearly transverse to quadrate, cylindrical; antennal clubs distinct and formed of apical 2-3 antennomeres; metasternal length between meso- and metacoxae one and a half or more length of mesocoxae … 8
8. (7) Lateral antebasal foveae not visible in dorsal view; elytra with discal striae slightly convergent with sutural striae toward apex, lacking sutural foveae at base … Vasse Chandler
Lateral antebasal foveae visible in dorsal view; elytra with discal striae curving away from sutural stria near apex, sutural foveae distinct … Antipodebaxis Chandler
9. (6) Head prolonged apically, lacking ventrolateral carinae beneath eyes, ventrolateral margins broadly concave to rounded; frontal fovea large and distinct or barely indicated; elytra with 3 basal foveae (2 in Startes from New Zealand) … 10
Head triangular to transverse, not prolonged apically, often with ventrolateral carinae, frontal fovea usually lacking (present in some Reichenbachia); elytra with 0-2 basal foveae (3 basal foveae in some Reichenbachia and Mundaring) … 11
10. (9) Pronotum with sharply defined antebasal sulcus nearly straight, median antebasal fovea barely wider than sulcus; fourth segments of maxillary palpi elongate, more than three times as long as wide, slightly narrowed at base … Paluma Chandler
Pronotum with antebasal sulcus distinct to lacking, medially angulate or arcuate when present, median antebasal fovea large, as large as lateral antebasal foveae; fourth segments of maxillary palpi shorter, less than three times as long as wide … Startes Broun
11. (9) Head with narrow postantennal slits originating at lateral margins, ventrolateral margins sharply rounded beneath eyes; pronotum with lateral antebasal foveae large and setose, median antebasal fovea small and nude or lacking … Wiangaree Chandler
Head lacking postantennal slits at lateral margins; ventrolateral margins of head more broadly rounded or carinate; pronotal pattern of foveation variable … 12
12. (11) Antennomeres IX wide and very thin, appressed to antennomeres X, apical two antennomeres subequal in size, forming large clubs; tibiae and femora strongly flattened … Malleecola Oke
Antennomeres IX not nearly so wide and transverse, antennomeres X clearly smaller than eleventh; tibiae cylindrical, femora not strongly flattened … 13
13. (12) Abdomen with lateral margins of abdomen mostly broadly rounded, paratergites reduced to single carinae, thin sulci, or lacking in apical half of segments … 14
Abdomen with lateral margins of first tergite angulate, paratergites clearly defined and nearly horizontal, though they may be quite narrow … 15
14. (13) Antennomeres X slightly larger and same form as IX; abdomen with first visible tergite (IV) shorter than remaining tergites combined … Eupinopsis Raffray
Antennomeres X clearly larger than IX, transverse; first visible tergite (IV) as long as to twice as long as rest of abdomen … Batraxis Raffray
15. (13) Head with ventrolateral carinae extending from base to beneath eyes and splitting to form outer margins of setose pits, in lateral view margins of head often angulate at area of pits … 16
Head with ventrolateral margins smoothly rounded or carinate, but lacking setose pits and not strongly angulate beneath eyes … 17
16. (15) Each elytron with two basal foveae; pronotum with distinct basal band exhibiting a series of short, evenly spaced longitudinal carinae, band not visible in heavily punctate species … Woodenbong Chandler
Elytra lacking basal foveae; pronotum with basal band narrower, less distinct … Wollomombi Chandler
17. (15) Pronotum with median antebasal fovea present, may be small, or pronotal disc at base impressed or excavate (one species from Western Australia with median fovea variably present); pronotal base often with transverse roughened or coarsely punctate band, or narrowly constricted … 18
Pronotum lacking median antebasal fovea, basal portion of disc convex; texture at base similar to center of pronotal disc … 24
18. (17) Tarsal claws long, longer than adjacent third tarsomeres; head in lateral view robust, as long as deep … Pedisinops Newton and Chandler
Tarsal claws shorter than adjacent third tarsomeres, usually half as long or less; head longer than depth in lateral view … 19
19. (18) Pronotum with setose lateral antebasal foveae … 20
Pronotum with nude lateral antebasal foveae … 21
20. (19) Elytra lacking basal foveae and discal striae; median gular carina prominent, abruptly curving to gular foveae in lateral view … Bundjulung Chandler
Elytra with basal foveae and discal striae; median gular carina low in lateral view … Reichenbachia Leach
21. (19) Pronotum with median and lateral antebasal foveae subequal in size; second visible sternite (IV) with basolateral sulci deep and densely setose to base shallowly depressed and nearly glabrous, line of posteriorly-directed setae from first visible sternite (III) over sulcus/depression distinct … Eupinolus Oke
Pronotum with lateral antebasal foveae smaller than median antebasal fovea or lateral foveae lacking; base of second visible sternite (IV) flat, not densely setose, lacking line of posteriorly-directed setae from apex of first visible sternite (III) … 22
22. (21) Head with prominent frontal rostrum; pronotal base same texture as disc, often broadly impressed at middle with basolateral angles of pronotum sharply angulate … Wataranka Chandler
Head with frontal rostrum low; pronotal base with poorly defined band of coarser punctures … 23
23. (22) Head with median gular carina prominent in lateral view; abdomen nearly as long as elytra in dorsal view … Storeyella Chandler
Head with median gular carina low, not obvious in lateral view; first visible tergite (IV) less than half as long as elytra in dorsal view, remaining tergites not or poorly visible … Iluka Chandler
24. (17) Each elytron with three basal foveae; median gular carina prominent, abruptly curving to gular foveae, ventrolateral margins beneath eyes concave; first visible tergite (IV) elongate, nearly as long as elytra … Mundaring Chandler
Each elytron with 0-2 basal foveae; median gular carina low, as prominent as ventrolateral carinae when they are present, ventrolateral margins beneath eyes convex or carinate; first visible tergite (IV) no more than two-thirds elytral length … 25
25. (24) Pronotum with lateral antebasal foveae and vertexal foveae distinct and setose … Briara Reitter
Pronotum with lateral antebasal foveae and vertexal foveae small and nude, lateral antebasal pronotal foveae often punctiform to lacking … 26
26. (25) Elytra lacking discal striae or basal foveae … Eupines King
Elytra with discal striae extending to near apices, basal foveae small or punctiform … 27
27. (26) Head with width of front separating first antennomeres narrow, no wider than width of first antennomeres; pronotum with lateral antebasal foveae distinct … Eupinella Raffray
Head with width of front separating first antennomeres wider, width clearly greater than width of first antennomeres; pronotum with lateral antebasal foveae punctiform to lacking … Anarmoxys Raffray
Diagnosis References
Chandler, D.S. 2001. Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Gainesville, FL : Memoirs on Entomology, International Vol. 15 viii + 560 pp. [Date published 30 April 2001] [290]
Raffray, A. 1908. Coleoptera fam. Pselaphidae. 1-487, pls i-ix in Wytsman, P. (ed.). Genera Insectorum, 64th fascicule. Rome : Wytsman. [190]
General References
Chandler, D.S. 2001. Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Gainesville, FL : Memoirs on Entomology, International Vol. 15 viii + 560 pp. [Date published 30 April 2001]
History of changes
| Published | As part of group | Action Date | Action Type | Compiler(s) |
|---|---|---|---|---|
| 01-Jul-2020 | 21-Dec-2011 | MODIFIED | ||
| 12-Feb-2010 | (import) |