Museums
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Sub-publications- Boucomont, A. & Gillet, J.J.E. 1927. Scarabaeidae: Coprinae II. pp. 101-264 in Schenkling, S. (ed.). Coleopterorum Catalogus Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 90.
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