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Ward, R.D. 1979. Metathoracic wing structures as phylogenetic indicators in the Adephaga (Coleoptera). pp. 181-191
in
Erwin, T.L., Ball, G.E., Whitehead, D.R. & Halpern, A.L. (eds).
Carabid beetles: Their evolution, natural history, and classification
. The Hague : W. Junk vii 635 pp.
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