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Walter, G.H. & Zalucki, M.P. 1999. Rare Butterflies and Theories of Evolution and Ecology. pp. 349-368
in
Kitching, R.L., Scheermeyer, E., Jones, R.L. & Pierce, N.E. (eds).
Biology of Australian Butterflies
. Collingwood : CSIRO Publishing xvi 395 pp.
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