Superfamily PTEROPHOROIDEA

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Family MACROPIRATIDAE

Introduction

[After Shaffer 1996]
The family was erected by Meyrick in 1932 for his species Macropiratis halieutica. An older name, Agdistopis Hampson, 1917, was originally described in the Pyralidae, subfamily Galleriinae. Whalley (1964a) synonymised the genera and transferred them to the Pterophoridae. Macropiratidae are known from Australia from the single species Agdistopis halieutica (Meyrick, 1932), described from Fiji and extending west to New Guinea and Australia. The genus contains two further species, sinhala Fletcher from Asia and griveaudi Gibeaux from Madagascar.

Many specialists did not accept Agdistopis as belonging to the Pterophoridae as the species lack the erect modified scales on the underside of the hindwing. Minet (1991) assigned the group as the Macropiratinae to the Pterophoridae, even though, as he stated, it differs from the remaining pterophorids in lacking two important derived features. Gielis (1993) has, however, treated the group with family status within the Pterophoroidea, where it is retained here. Externally Agdistopis bears a striking resemblance to the true pterphorid genus Agdistis, a genus not represented in Australia, and can only be satisfactorily distinguished by dissection of the genitalia and by the lack of the modified scales of the hindwing.

The single foodplant record known comes from a specimen from the Northern Territory where an adult emerged from fruit of Secamone elliptica. The moth is also recorded from Queensland.

Further studies are needed on this genus, dissections may well indicate that there are additional species mixed in with both the sinhala and the halieutica series.

 

General References

Gielis, C. 1993. Generic revision of the superfamily Pterophoroidea (Lepidoptera). Zoologische Verhandelingen (Leiden) 290: 1-139 [Reprinted and published by the author, Amsterdam, 1994]

Minet, J. 1991. Tentative reconstruction of the ditrysian phylogeny (Lepidoptera: Glossata). Entomologica Scandinavica 22: 69-95

Shaffer, M. 1996. Macropiratidae. p. 156 in Nielsen, E.S., Edwards, E.D. & Rangsi, T.V. (eds). Checklist of the Lepidoptera of Australia. Monographs on Australian Lepidoptera. Collingwood : CSIRO Publishing Vol. 4 xiv 529 pp. & CD-ROM.

Whalley, P.E.S. 1964. Catalogue of the world genera of the Thyrididae (Lepidoptera) with type selection and syntypes. Annals and Magazine of Natural History 13 7: 115-127

 

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Genus Agdistopis Hampson, 1917

 

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Species Agdistopis halieutica (Meyrick, 1932)

 

Generic Combinations

 

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Family PTEROPHORIDAE

Introduction

[After Shaffer & Nielsen 1996]
The first Pterophoridae were described from Australia by Walker in 1864, but the oldest described species listed were named from abroad and have subsequently been found in Australia. However, the oldest name in the Australian list is an introduced species, Wheeleria spilodactylus (Curtis, 1827). In the 19th century, Newman, Meyrick and Lucas added several species to the Australian fauna and several more were added in the 20th century, as recently as 1991. Today 49 species are recognised as occurring in Australia. The world fauna of the family consists of over 1200 described species. Further research will reveal additional species as several undescribed species are known.

The family was originally described by Latreille in 1802 and three subfamilies are currently known from Australia. Common (1990) and Nielsen & Common (1991) recognised Agdistinae, Platyptiliinae and Pterophorinae. Research since then has shown that true Agdistinae do not occur in Australia, and that the Platyptiliinae are synonymous with Pterophorinae (Gielis 1993). The subfamilies Ochyroticinae, Deuterocopinae and Pterophorinae are currently recognised in the Australian fauna. Besides the confusion over the placement of the genus Alucita (see ICZN, 1957), the group as a whole has always been recognised correctly and not confused with other families, except for the inclusion of the Maropiratidae.

Foodplant records are: Deuterocopus triannulatus on Vitis lanata; Stenoptilia zophodactylus on flowers of Gentianella diemensis; Platyptilia omissalis on Parahebe perfoliata; Sphenarches anisodactylus on flower buds and flowers of Pelaronium, also known to occur abroad on Dolichos lablab and on Lagenaria; Stangeia xerodes on foliage of Acacia and Imbophorus aptalis on undersides of leaves of Astrotricha floccosa.

The Ochyroticinae and Agdistinae are easily recognised with both pairs of wings undivided. In the Deuterocopinae the forewings have two clefts and the hindwings likewise, whereas the Pterophorinae have a single cleft, of varying depth in the forewing and two deep clefts in the hindwing. All true Pterophoridae have a line of erect modified scales on the underside of the hindwing.

 

General References

Common, I.F.B. 1990. Moths of Australia. Carlton : Melbourne University Press vi+535 pp., 32 pls.

Gielis, C. 1993. Generic revision of the superfamily Pterophoroidea (Lepidoptera). Zoologische Verhandelingen (Leiden) 290: 1-139 [Reprinted and published by the author, Amsterdam, 1994]

Nielsen, E.S. & Common, I.F.B. 1991. Lepidoptera (Moths and Butterflies). pp. 817-915, pls 7, 8 in CSIRO (ed.). The Insects of Australia. 2 vols. Carlton : Melbourne University Press xiii + vi + 1075 pp., 8 pls.

Shaffer, M. 1996. Pterophoridae. pp. 157-158 in Nielsen, E.S., Edwards, E.D. & Rangsi, T.V. (eds). Checklist of the Lepidoptera of Australia. Monographs on Australian Lepidoptera. Collingwood : CSIRO Publishing Vol. 4 xiv 529 pp. & CD-ROM.

 

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Subfamily DEUTEROCOPINAE

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Genus Deuterocopus Zeller, 1852

 

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Species Deuterocopus atrapex Fletcher, 1910

 

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Species Deuterocopus honoratus Meyrick, 1921

 

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Species Deuterocopus socotranus Rebel, 1907

 

Ecological Descriptors

Herbivore (associated flora: Vitis lanata Roxb. [VITACEAE] grape (recorded as foodplant for Deuterocopus triannulatus)).

 

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Species Deuterocopus tengstroemi Zeller, 1852

 

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Genus Hexadactilia Fletcher, 1910

 

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Species Hexadactilia civilis Meyrick, 1921

 

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Species Hexadactilia trilobata Fletcher, 1910

 

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Subfamily OCHYROTICINAE

 

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Genus Ochyrotica Walsingham, 1891

 

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Species Ochyrotica kurandica Arenberger, 1988

 

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Subfamily PTEROPHORINAE

 

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Genus Bipunctiphorus Gibeaux, 1994

 

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Species Bipunctiphorus euctimena (Turner, 1913)

 

Generic Combinations

 

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Genus Cosmoclostis Meyrick, 1886

 

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Species Cosmoclostis aglaodesma Meyrick, 1886

 

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Species Cosmoclostis hemiadelpha Fletcher, 1947

 

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Genus Crombrugghia Tutt, 1906

 

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Species Crombrugghia wahlbergi (Zeller, 1851)

 

Generic Combinations

 

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Genus Hellinsia Tutt, 1905

 

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Species Hellinsia balanotes (Meyrick, 1908)

Groundsel Plume Moth

 

Generic Combinations

 

Distribution

Extra Distribution Information

Established, see Julien 1992: 10

Introduced from USA.


Distribution References

Ecological Descriptors

Herbivore (associated flora: Baccharis halimifolia L. [ASTERACEAE] Groundsel).

 

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Species Hellinsia lienigianus (Zeller, 1852)

 

Generic Combinations

 

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Species Hellinsia tinctidactylus (Newman, 1856)

 

Generic Combinations

 

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Genus Hepalastis Gibeaux, 1994

 

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Species Hepalastis pumilio (Zeller, 1873)

 

Generic Combinations

 

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Genus Imbophorus Arenberger, 1991

 

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Species Imbophorus aptalis (Walker, 1864)

 

Generic Combinations

 

Ecological Descriptors

Herbivore (associated flora: Astrotricha floccosa DC. [ARALIACEAE] (on undersides of leaves)).

 

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Species Imbophorus leucophasma (Turner, 1911)

 

Generic Combinations

 

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Species Imbophorus pallidus Arenberger, 1991

 

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Genus Lantanophaga Zimmerman, 1958

 

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Species Lantanophaga pusillidactylus (Walker, 1864)

Lantana Plume Moth

 

Generic Combinations

 

Distribution

Extra Distribution Information

Established, see (Julien, M.H., 1992) [138].

Introduced from ?Mexico.


Distribution References

Ecological Descriptors

Herbivore (associated flora: Lantana [VERBENACEAE]).

 

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Genus Megalorhipida Amsel, 1935

 

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Species Megalorhipida defectalis (Walker, 1864)

 

Generic Combinations

 

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Species Megalorhipida leucodactyla (Fabricius, 1794)

 

Generic Combinations

 

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Genus Nippoptilia Matsumura, 1931

 

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Species Nippoptilia cinctipedalis (Walker, 1864)

 

Generic Combinations

 

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Genus Oxyptilus Zeller, 1841

 

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Species Oxyptilus regulus Meyrick, 1906

 

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Genus Platyptilia Hübner, 1825

 

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Species Platyptilia celidotus (Meyrick, 1885)

 

Generic Combinations

 

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Species Platyptilia isodactylus (Zeller, 1852)

 

Taxonomic Decision for Synonymy

 

Generic Combinations

 

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Species Platyptilia omissalis Fletcher, 1926

 

Ecological Descriptors

Herbivore (associated flora: Parahebe perfoliata (R.Br.) B.G.Briggs & Ehrend. [SCROPHULARIACEAE] Blue Veronica).

 

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Genus Pterophorus Schaffer, 1766

 

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Species Pterophorus albidus (Zeller, 1852)

 

Generic Combinations

 

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Species Pterophorus lacteipennis (Walker, 1864)

 

Generic Combinations

 

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Genus Sinpunctiptilia Arenberger, 2006

 

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Species Sinpunctiptilia emissalis (Walker, 1864)

 

Generic Combinations

 

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Species Sinpunctiptilia tasmaniae Arenberger, 2006

 

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Genus Sphenarches Meyrick, 1886

 

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Species Sphenarches anisodactylus (Walker, 1864)

 

Generic Combinations

 

Ecological Descriptors

Florivore (associated flora: Pelargonium [GERANIACEAE]).

Extra Ecological Information

On flowers and buds

 

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Species Sphenarches zanclistes (Meyrick, 1905)

 

Generic Combinations

 

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Genus Stangeia Tutt, 1906

 

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Species Stangeia xerodes (Meyrick, 1886)

 

Generic Combinations

 

Ecological Descriptors

Herbivore (associated flora: Acacia [FABACEAE] Wattle).

 

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Genus Stenodacma Amsel, 1959

 

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Species Stenodacma pyrrhodes (Meyrick, 1890)

 

Generic Combinations

 

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Genus Stenoptilia Hübner, 1825

 

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Species Stenoptilia leuconephes (Meyrick, 1886)

 

Generic Combinations

 

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Species Stenoptilia phaeonephes (Meyrick, 1886)

 

Generic Combinations

 

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Species Stenoptilia zophodactylus (Duponchel, 1840)

 

Generic Combinations

 

Ecological Descriptors

Florivore (associated flora: Gentianella diemensis (Griseb.) J.H.Willis [GENTIANACEAE] Native Gentian).

 

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Genus Stenoptilodes Zimmerman, 1958

 

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Species Stenoptilodes taprobanes (R. Felder & Rogenhofer, 1875)

 

Generic Combinations

 

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Genus Tetraschalis Meyrick, 1887

 

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Species Tetraschalis arachnodes Meyrick, 1887

 

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Genus Tomotilus Yano, 1961

 

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Species Tomotilus saitoi Yano, 1961

 

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Genus Trichoptilus Walsingham, 1880

 

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Species Trichoptilus ceramodes Meyrick, 1886

 

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Species Trichoptilus inclitus T.P. Lucas, 1892

 

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Species Trichoptilus scythrodes Meyrick, 1886

 

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Genus Wheeleria Tutt, 1905

 

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Species Wheeleria spilodactylus (Curtis, 1827)

 

Generic Combinations

 

Distribution

Extra Distribution Information

Established, J. Weiss, pers. comm.

Introduced from Spain.


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Genus Xyroptila Meyrick, 1908

 

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11-May-2011 11-May-2011 MODIFIED
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Species Xyroptila elegans Kovtunovich & Ustjuzhanin, 2006

 

History of changes

Note that this list may be incomplete for dates prior to September 2013.
Published As part of group Action Date Action Type Compiler(s)
22-Jan-2011 ADDED

Species Xyroptila kuranda Kovtunovich & Ustjuzhanin, 2006

 

History of changes

Note that this list may be incomplete for dates prior to September 2013.
Published As part of group Action Date Action Type Compiler(s)
22-Jan-2011 ADDED

Species Xyroptila marmarias Meyrick, 1908

 

History of changes

Note that this list may be incomplete for dates prior to September 2013.
Published As part of group Action Date Action Type Compiler(s)
11-May-2011 11-May-2011 MODIFIED
12-Feb-2010 (import)

Species Xyroptila peltastes (Meyrick, 1908)

 

Generic Combinations

 

History of changes

Note that this list may be incomplete for dates prior to September 2013.
Published As part of group Action Date Action Type Compiler(s)
11-May-2011 11-May-2011 MODIFIED
12-Feb-2010 (import)

Species Xyroptila uluru Kovtunovich & Ustjuzhanin, 2006

 

History of changes

Note that this list may be incomplete for dates prior to September 2013.
Published As part of group Action Date Action Type Compiler(s)
22-Jan-2011 ADDED