Family PASSALIDAE

Compiler and date details

1 December 2002 - Gerasimos Cassis & Tom A. Weir; updated by Andrew A. Calder, CSIRO Entomology, Canberra, Australia

Introduction

The Passalidae are a cosmopolitan family of Scarabaeoidea, comprising about 27 genera and 500 species. They occur mostly in tropical and subtropical regions (Lawrence 1982). Hincks & Dibb (1958) listed seven subfamilies, but recent work by Reyes-Castillo (1970) reduced this to two subfamilies, the Aulacocyclinae and the Passalinae. The classification followed in this catalogue is that of Reyes-Castillo (1970).

Percheron (1835) and Kaup (1871) were the first to monograph the world fauna. Gravely (1914, 1918) revised the world fauna, provided the first modern treatment, discussed the classification, morphology, distribution, and provided new synonymies and checklists. Masters (1886) catalogued the Australian fauna, Kuwert (1891, 1898) described 18 species for Australia and Dibb (1938) reviewed the Australian fauna although he added no new species. Hincks & Dibb (1935) catalogued the world fauna and provided a supplement in 1958.

The Australian fauna comprises nine genera and 35 species; the largest number of species are assigned to Aulacocyclus Kaup, and the rest of the fauna to the Passalinae. Other large genera include Mastachilus Kaup and Pharochilus Kaup, and most of the Australian species are found in the north and the east of the continent.

BIOLOGY
This group has specialised biology and exhibits sub-social behaviour. Passalids tend to form colonies in rotten logs and the adults protect the young. The larvae feed on wood pulp prepared by adults and there is communication between larvae and adults by stridulation (Matthews 1984). In the adults, stridulation is achieved by friction between the wings and upper surface of the abdomen. The larvae produce a high pitched sound by scraping the reduced hind leg over a file on the midcoxa (Lawrence & Britton 1991).

Diagnosis

The adults of this family are usually black, large in size (20–60 mm) and somewhat flattened, with the head prognathous and sometimes armed with a short horn. The antennae are characteristically curved and there is a narrow pedicel between the prothorax and the elytra. The scutellum is not visible and the elytral striae are conspicuous (Lawrence & Britton 1991). The larvae are distinguished by the mandibles without a ventral process, the antennae are two segmented, and the body shape is less curved than other scarabaeoid larvae. The hind legs are greatly reduced and unsegmented (Ritcher 1966).

Diagnosis References

Lawrence, J.F. & Britton, E.B. 1991. Chapter 35. Coleoptera (Beetles). pp. 543-683 in Division of Entomology, CSIRO (ed.). The Insects of Australia. Ithaca, New York : Cornell University Press Vol. 2.

Ritcher, P.O. 1966. White grubs and their allies. A study of North American scarabaeoid larvae. Oregon State Monographs. Studies in Entomology 4: 1-219

General References

Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103-124

Gravely, F.H. 1914. An account of the Oriental Passalidae (Coleoptera) based primarily on the collection in the Indian Museum. Memoirs of the Indian Museum 3: 177-353

Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142.

Hincks, W.D. & Dibb, J.R. 1958. Passalidae. pp. 1-32 in Hincks, W.D. (ed.). Coleopterorum Catalogus Supplementa. Gravenhage : W. Junk Pt 142.

Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125

Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192

Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349

Lawrence, J.F. 1982. Coleoptera. pp. 482-553 in Parker, S.P. (ed.). Synopsis and Classification of Living Organisms. New York : McGraw Hill Vol. 2 vii 1232 pp.

Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Lucanidae and Scarabaeidae. Proceedings of the Linnean Society of New South Wales 2 1: 21-126

Matthews, E.G. 1984. A Guide to the Genera of Beetles of South Australia. Part 3. Polyphaga: Eucinetoidea, Dascilloidea and Scarabaeoidea. Adelaide : South Australian Museum 60 pp.

Percheron, A.R. 1835. Monographie des Passales et des genres qui en ont été séparés; accompagnée de planches dessinées par l'auteur, ou toutes les espèces ont été figurées. Paris : Mercklein iii 107 pp. 7 pls.

Reyes-Castillo, P. 1970. Coleoptera, Passalidae: morfología y división en grandes grupos; géneros americanos. Folia Entomologica Mexicana 1970(20–22): 1-240

Ritcher, P.O. 1966. White grubs and their allies. A study of North American scarabaeoid larvae. Oregon State Monographs. Studies in Entomology 4: 1-219

History of changes

Note that this list may be incomplete for dates prior to September 2013.

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		12-Feb-2010	(import)

Subfamily Aulacocyclinae

History of changes

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Genus Aulacocyclus Kaup, 1868

Aulacocyclus Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [4].
Type species:
Passalus edentulus W.S. Macleay, 1826 by subsequent designation, see Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [8] (considered as the first fixation of the type by [22]).
Caulifer Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [22].
Type species:
Caulifer macleayi Kaup, 1871 by monotypy.

Taxonomic Decision for Synonymy

Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [13] (also for extralimital synonymy)

Distribution

States

New South Wales, Queensland, Victoria

Extra Distribution Information

Oriental Region.

Note that conversion of the original AFD map of states, drainage basins and coastal and oceanic zones to IBRA and IMCRA regions may have produced errors. The new maps will be reviewed and corrected as updates occur. The maps may not indicate the entire distribution. See further details below.

IBRA

Australian Alps (AA), Arnhem Coast (ARC), Arnhem Plateau (ARP), Avon Wheatbelt (AW), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Ben Lomond (BEL), Broken Hill Complex (BHC), Burt Plain (BRT), Central Arnhem (CA), Carnarvon (CAR), Channel Country (CHC), Central Kimberley (CK), Central Mackay Coast (CMC), Coolgardie (COO), Cobar Peneplain (CP), Central Ranges (CR), Cape York Peninsula (CYP), Daly Basin (DAB), Darwin Coastal (DAC), Desert Uplands (DEU), Dampierland (DL), Davenport Murchison Ranges (DMR), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Esperance Plains (ESP), Eyre Yorke Block (EYB), Finke (FIN), Flinders Lofty Block (FLB), Flinders (FLI), Gascoyne (GAS), Gawler (GAW), Gibson Desert (GD), Gulf Fall and Uplands (GFU), Geraldton Sandplains (GS), Great Sandy Desert (GSD), Gulf Coastal (GUC), Gulf Plains (GUP), Great Victoria Desert (GVD), Hampton (HAM), Jarrah Forest (JF), Kanmantoo (KAN), King (KIN), Little Sandy Desert (LSD), MacDonnell Ranges (MAC), Mallee (MAL), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Murchison (MUR), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), Northern Kimberley (NK), NSW North Coast (NNC), NSW South Western Slopes (NSS), Nullarbor (NUL), Ord Victoria Plain (OVP), Pine Creek (PCK), Pilbara (PIL), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Stony Plains (STP), Sturt Plateau (STU), Swan Coastal Plain (SWA), Tanami (TAN), Tasmanian Central Highlands (TCH), Tiwi Cobourg (TIW), Tasmanian Northern Midlands (TNM), Tasmanian Northern Slopes (TNS), Tasmanian South East (TSE), Tasmanian Southern Ranges (TSR), Tasmanian West (TWE), Victoria Bonaparte (VB), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Warren (WAR), Wet Tropics (WT), Yalgoo (YAL) ; NSW, Qld, Vic: Australian Alps (AA), Arnhem Coast (ARC), Arnhem Plateau (ARP), Avon Wheatbelt (AW), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Ben Lomond (BEL), Broken Hill Complex (BHC), Burt Plain (BRT), Central Arnhem (CA), Carnarvon (CAR), Channel Country (CHC), Central Kimberley (CK), Central Mackay Coast (CMC), Coolgardie (COO), Cobar Peneplain (CP), Central Ranges (CR), Cape York Peninsula (CYP), Daly Basin (DAB), Darwin Coastal (DAC), Desert Uplands (DEU), Dampierland (DL), Davenport Murchison Ranges (DMR), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Esperance Plains (ESP), Eyre Yorke Block (EYB), Finke (FIN), Flinders Lofty Block (FLB), Flinders (FLI), Gascoyne (GAS), Gawler (GAW), Gibson Desert (GD), Gulf Fall and Uplands (GFU), Geraldton Sandplains (GS), Great Sandy Desert (GSD), Gulf Coastal (GUC), Gulf Plains (GUP), Great Victoria Desert (GVD), Hampton (HAM), Jarrah Forest (JF), Kanmantoo (KAN), King (KIN), Little Sandy Desert (LSD), MacDonnell Ranges (MAC), Mallee (MAL), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Murchison (MUR), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), Northern Kimberley (NK), NSW North Coast (NNC), NSW South Western Slopes (NSS), Nullarbor (NUL), Ord Victoria Plain (OVP), Pine Creek (PCK), Pilbara (PIL), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Stony Plains (STP), Sturt Plateau (STU), Swan Coastal Plain (SWA), Tanami (TAN), Tasmanian Central Highlands (TCH), Tiwi Cobourg (TIW), Tasmanian Northern Midlands (TNM), Tasmanian Northern Slopes (TNS), Tasmanian South East (TSE), Tasmanian Southern Ranges (TSR), Tasmanian West (TWE), Victoria Bonaparte (VB), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Warren (WAR), Wet Tropics (WT), Yalgoo (YAL)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: Murray-Darling basin, SE coastal
- Queensland: Lake Eyre basin, Murray-Darling basin, NE coastal
- Victoria: SE coastal
New Caledonia
Papua New Guinea

Distribution References

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [8]

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus aliicornis Kuwert, 1897

Aulacocyclus aliicornis Kuwert, A.F. 1897. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 4: 274-306 [277].
Type data:
Holotype MNHP, Australia (as Australien).
Aulacocyclus aliicornis sulcatipons Kuwert, A.F. 1897. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 4: 274-306 [279].
Type data:
Holotype MNHP, Australia (as Australien).

Taxonomic Decision for Synonymy

Gravely, F.H. 1914. An account of the Oriental Passalidae (Coleoptera) based primarily on the collection in the Indian Museum. Memoirs of the Indian Museum 3: 177-353 [273]

Distribution

States

Queensland

IBRA

Qld: Brigalow Belt North (BBN), Brigalow Belt South (BBS), Central Mackay Coast (CMC), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Mulga Lands (ML), Nandewar (NAN), NSW North Coast (NNC), South Eastern Queensland (SEQ), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- Queensland: Murray-Darling basin, NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus collaris Blackburn, 1896

Aulacocyclus collaris Blackburn, T. 1896. Further notes on Australian Coleoptera, with descriptions of new genera and species. Part XX. Transactions of the Royal Society of South Australia 20: 233-259 [233].
Type data:

Syntype(s) BMNH, Australia; SAMA I33 1 specimen, Australia.

Distribution

States

Queensland

IBRA

Qld: Brigalow Belt North (BBN), Brigalow Belt South (BBS), Central Mackay Coast (CMC), Cape York Peninsula (CYP), Desert Uplands (DEU), Einasleigh Uplands (EIU), NSW North Coast (NNC), South Eastern Queensland (SEQ), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus deyrolli Kaup, 1868

Aulacocyclus deyrolli Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [7].
Type data:
Syntype(s) 3 specimens, St David, Réunion Is. (as St. Denis auf Réunion).

Distribution

States

Queensland, Victoria

Extra Distribution Information

Réunion Is.

IBRA

Qld, Vic: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Central Mackay Coast (CMC), Cape York Peninsula (CYP), Desert Uplands (DEU), Einasleigh Uplands (EIU), Flinders (FLI), Murray Darling Depression (MDD), Naracoorte Coastal Plain (NCP), NSW North Coast (NNC), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal
- Victoria: SE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus edentulus (W.S. Macleay, 1826)

Passalus edentulus Macleay, W.S. 1826. Annulosa. Catalogue of insects collected by Captain King, R.N. pp. 438-457 in King, P.P. (ed.). Narrative of a Survey of the Intertropical and Western Coasts of Australia. Performed between the years 1818 and 1822; with Appendix B. London : John Murray Vol. 2 viii 637 pp., 9 pls. [Date published 15 Apr 1826: imprint 1827] [publication date 15 April 1826 Common, I.F.B. & Moulds, M.S. 1973. The date of publication of Captain King's narrative of a survey of the intertropical and western coasts of Australia. Journal of the Society for the Bibliography of Natural History 6: 257–259 ("an initial small issue was published in the first half of 1826 and a second much larger issue in 1827")] [439].
Type data:
Holotype ANIC(MMUS), Australia, no exact type locality specified.
Passalus furcicornis Boisduval, J.A. 1835. Faune Entomologique de l'Océan Pacifique, avec l'illustration des insects nouveaux recueillis pendent de voyage. 2me Partie. Coléoptères et autres Ordres. In, D'Urville, D., César, J. S. (ed.). Voyage de Découvertes de l'Astrolabe exécuté par ordre du Roi, pendant les annés 1826–1829, sous le commandement de M.J. Dumont d'Urville. Paris : J. Tastu Vol. 7 716 pp. [242].
Type data:
Status unknown, whereabouts unknown, Port Jackson (as Port-Jakson), NSW.
Passalus cylindraceus Percheron, A.R. 1835. Monographie des Passales et des genres qui en ont été séparés; accompagnée de planches dessinées par l'auteur, ou toutes les espèces ont été figurées. Paris : Mercklein iii 107 pp. 7 pls. [103].
Type data:
Syntype(s) whereabouts unknown, ?Australia.
Secondary source:
Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [9].
Aulacocyclus terioides Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [170].
Type data:
Holotype MNHP, Australia (as Neuholland).

Taxonomic Decision for Synonymy

Gravely, F.H. 1914. An account of the Oriental Passalidae (Coleoptera) based primarily on the collection in the Indian Museum. Memoirs of the Indian Museum 3: 177-353 [275]

Distribution

States

New South Wales, Queensland, Victoria

IBRA

NSW, Qld, Vic: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Broken Hill Complex (BHC), Central Mackay Coast (CMC), Cobar Peneplain (CP), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Flinders (FLI), Murray Darling Depression (MDD), Mulga Lands (ML), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), NSW North Coast (NNC), NSW South Western Slopes (NSS), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: Murray-Darling basin, SE coastal
- Queensland: Murray-Darling basin, NE coastal
- Victoria

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Mesa, A., Ferreira, A. & Martins, U. 1978. The chromosomes of an Australian passalid, Aulacocylcus edentulus Macl. (Coleoptera, Passalidae, Eulacocyclinae). Journal of the Australian Entomological Society 17: 385-388 (chromosome number)

Schuster, J.C. & Reyes-Castillo, P. 1981. New World genera of Passalidae (Coloptera): a revision of larvae. Anales de la Escuela Nacional de Ciencias Biologicas, Mexico 25: 79-116 (taxonomy)

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus fracticornis Kuwert, 1891

Aulacocyclus fracticornis Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [170] [erroneously considered as a nom. nud. by Dibb, J.R. 1935. Further notes on Aulacocyclus (Col., Passalidae). Stylops 4: 227–233].
Type data:
Status unknown, MNHP, South Seas (as Südsee (Patria?), no exact locality specified.
Type locality references:
Dibb, J.R. 1932. The Australian species of the genus Aulacocyclus (Col., Passalidae). Stylops 1: 257-263 [258].
Secondary source:
Dibb, J.R. 1935. Further notes on Aulacocyclus (Col., Passalidae). Stylops 4: 227-233.
Aulacocyclus rotundatoclypeatus Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [170] [erroneously considered as a nom. nud. by Dibb, J.R. 1935. Further notes on Aulacocyclus (Col., Passalidae). Stylops 4: 227–233].
Type data:
Holotype MNHP, Cape York (as Cap York), QLD.
Secondary source:
Dibb, J.R. 1935. Further notes on Aulacocyclus (Col., Passalidae). Stylops 4: 227-233.
Aulacocyclus errans Blackburn, T. 1896. Further notes on Australian Coleoptera, with descriptions of new genera and species. Part XX. Transactions of the Royal Society of South Australia 20: 233-259 [233].
Type data:
Syntype(s) BMNH, Australia.

Taxonomic Decision for Synonymy

Dibb, J.R. 1935. Further notes on Aulacocyclus (Col., Passalidae). Stylops 4: 227-233 [227] (also for extralimital synonymy)
Cassis, G. & Weir, T. A. 1992. Passalidae. pp. 20-30 in Houston, W.W.K. (ed.). Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Canberra : AGPS Vol. 9 544 pp. [22] (stat. nov.; synonymy of Dibb (1935), based on redescriptions by Kuwert, A.F. 1897. Die Passaliden dichotomisch bearbeitet. Novit. Zool. 4: 274–306[278], accepted here, although date priority is applied, thus A. fracticornis is considered the valid name)

Distribution

States

New South Wales, Queensland

IBRA

NSW, Qld: Brigalow Belt North (BBN), Brigalow Belt South (BBS), Central Mackay Coast (CMC), Cape York Peninsula (CYP), Desert Uplands (DEU), Einasleigh Uplands (EIU), NSW North Coast (NNC), Sydney Basin (SB), South East Corner (SEC), South Eastern Queensland (SEQ), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: SE coastal
- Queensland: NE coastal
Papua New Guinea

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus gravelyi Dibb, 1933

Aulacocyclus gravelyi Dibb, J.R. 1933. A new species of Aulacocyclus (Passalidae) from Australia. Entomologist's Monthly Magazine 69: 103-104 [104].
Type data:
Holotype MMUE, Australia, no exact type locality specified.
Paratype(s) MMUE.

Distribution

Extra Distribution Information

Exact locality unknown.

IBRA

Original AFD Distribution Data

Australian Region

Australia

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus hangayi Van Doesburg, 1992

Aulacocyclus hangayi Van Doesburg, P.H. 1992. A new species of Aulacocyclus from Australia (Coleoptera: Passalidae). Zoologische Mededelingen (Leiden) 66(29): 413-415 [413].
Type data:
Holotype AM K209194 ♀ (22.ix.1984), E of Middleton, W of Winton, QLD [22° 21' S 141° 33' E].

Distribution

States

Queensland

IBRA

Qld: Channel Country (CHC), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Simpson Strzelecki Dunefields (SSD)

Original AFD Distribution Data

Australian Region

Australia
- Queensland: Lake Eyre basin

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus kaupii Macleay, 1871

Aulacocyclus kaupii Macleay, W.J. 1871. Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales 2: 79-205 [173].
Type data:
Holotype AM, Gayndah, QLD.

Distribution

States

Queensland

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Froggatt, W.W. 1893. On the life-histories of Australian Coleoptera. Proceedings of the Linnean Society of New South Wales 2 8: 33-42 (biology)

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus macleayi (Kaup, 1871)

Caulifer macleayi Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [22].
Type data:
Holotype whereabouts unknown, Sydney (as Sidney), NSW.

Distribution

States

New South Wales

Extra Distribution Information

Known only from type locality.

IBRA

NSW: NSW North Coast (NNC), Sydney Basin (SB), South East Corner (SEC), South Eastern Queensland (SEQ)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: SE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus mastersii (Macleay, 1871)

Taeniocerus mastersii Macleay, W.J. 1871. Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales 2: 79-205 [174].
Type data:
Holotype AM, Gayndah, QLD.
Aulacocyclus foveipunctatus Kuwert, A.F. 1897. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 4: 274-306 [278].
Type data:
Holotype MNHP, Australia (as Australien).

Taxonomic Decision for Synonymy

Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103-124 [114]

Distribution

States

New South Wales, Queensland, Victoria

IBRA

NSW, Qld, Vic: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Central Mackay Coast (CMC), Cape York Peninsula (CYP), Desert Uplands (DEU), Einasleigh Uplands (EIU), Flinders (FLI), Murray Darling Depression (MDD), Naracoorte Coastal Plain (NCP), NSW North Coast (NNC), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: SE coastal
- Queensland: NE coastal
- Victoria: SE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

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Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Aulacocyclus rosenbergii Kaup, 1868

Aulacocyclus rosenbergii Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [7].
Type data:
Holotype Nias Is. (as Niasinsel), Indonesia.
Aulacocyclus percheroni Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [8].
Type data:

Syntype(s) 1 specimen, Brisbane, QLD (as Trisbane, Neuholland); 1 specimen, Brisbane, QLD (as Trisbane, Neuholland).
Secondary source:
Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [16].

Taxonomic Decision for Synonymy

Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [16]

Distribution

States

Queensland

Extra Distribution Information

Oriental Region.

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

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		12-Feb-2010	(import)

Species Aulacocyclus tambourinensis Mjöberg, 1917

Aulacocyclus tambourinensis Mjöberg, E. 1917. Results of Dr. E. Mjöberg's Swedish Scientific Expeditions to Australia 1910–1913. 14. Cetonidae, Rutelidae, Passalidae, Chrysomelidae: Subfam. Sagrinae, Cassidinae, Hispinae. Arkiv för Zoologi 11: 1-19 [10].
Type data:
Syntype(s) NHRM 18 specimens, Mt Tamborine, QLD.

Distribution

States

Queensland

Extra Distribution Information

Known only from type locality.

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

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		12-Feb-2010	(import)

Species Aulacocyclus teres (Percheron, 1841)

Passalus teres Percheron, A.R. 1841. Révision critique et supplément à la monographie du genre Passalus. Revue et Magasin de Zoologie (Paris) 2 3: 1-48 [39].
Type data:
Syntype(s) whereabouts unknown, ?Australia.
Secondary source:
Dibb, J.R. 1932. The Australian species of the genus Aulacocyclus (Col., Passalidae). Stylops 1: 257-263 [260].

Distribution

States

New South Wales, Queensland

IBRA

NSW, Qld: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Broken Hill Complex (BHC), Channel Country (CHC), Central Mackay Coast (CMC), Cobar Peneplain (CP), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Murray Darling Depression (MDD), Mulga Lands (ML), Nandewar (NAN), New England Tablelands (NET), NSW North Coast (NNC), NSW South Western Slopes (NSS), Riverina (RIV), Sydney Basin (SB), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

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		12-Feb-2010	(import)

Subfamily Passalinae

History of changes

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Genus Analaches Kuwert, 1891

Analaches Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [167].
Type species:
Analaches puberilis Kuwert, 1891 by subsequent designation, see Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [98].

Distribution

States

New South Wales, Queensland

IBRA

NSW, Qld: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Broken Hill Complex (BHC), Channel Country (CHC), Central Mackay Coast (CMC), Cobar Peneplain (CP), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Gulf Plains (GUP), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Nandewar (NAN), New England Tablelands (NET), NSW North Coast (NNC), NSW South Western Slopes (NSS), Riverina (RIV), Sydney Basin (SB), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales
- Queensland
Papua New Guinea

Distribution References

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [90]

History of changes

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Species Analaches australiensis (Stoliczka, 1873)

Cetejus australiensis Stoliczka, A. 1873. A contribution towards a monograph of the Indian Passalidae. Journal of the Asiatic Society of Bengal ns 42: 149-162 [157].
Type data:
Syntype(s) Australia (as Australien).
Secondary source:
Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349 [331].

Distribution

States

New South Wales, Queensland

IBRA

NSW, Qld: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Broken Hill Complex (BHC), Channel Country (CHC), Central Mackay Coast (CMC), Cobar Peneplain (CP), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Gulf Plains (GUP), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Nandewar (NAN), New England Tablelands (NET), NSW North Coast (NNC), NSW South Western Slopes (NSS), Riverina (RIV), Sydney Basin (SB), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales
- Queensland
Papua New Guinea

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

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		12-Feb-2010	(import)

Genus Austropassalus Mjöberg, 1917

Austropassalus Mjöberg, E. 1917. Results of Dr. E. Mjöberg's Swedish Scientific Expeditions to Australia 1910–1913. 14. Cetonidae, Rutelidae, Passalidae, Chrysomelidae: Subfam. Sagrinae, Cassidinae, Hispinae. Arkiv för Zoologi 11: 1-19 [11].
Type species:
Austropassalus hultgreni Mjöberg, 1917 by monotypy.
Notocetius Hincks, W.D. & Dibb, J.R. 1935. A new genus and species of Passalidae (Col.) from Australia. Entomologist's Monthly Magazine 71: 272-274 [272].
Type species:
Notocetius cornutus Hincks & Dibb, 1935 by monotypy.

Taxonomic Decision for Synonymy

Hincks, W.D. 1949. Systematic and synonymic notes on Passalidae (Col.). Annals and Magazine of Natural History 12 2: 56-64 [64]

Distribution

States

Queensland

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

History of changes

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Species Austropassalus hultgreni Mjöberg, 1917

Austropassalus hultgreni Mjöberg, E. 1917. Results of Dr. E. Mjöberg's Swedish Scientific Expeditions to Australia 1910–1913. 14. Cetonidae, Rutelidae, Passalidae, Chrysomelidae: Subfam. Sagrinae, Cassidinae, Hispinae. Arkiv för Zoologi 11: 1-19 [11].
Type data:
Syntype(s) NHRM 12 specimens, near Atherton, QLD.
Notocetius cornutus Hincks, W.D. & Dibb, J.R. 1935. A new genus and species of Passalidae (Col.) from Australia. Entomologist's Monthly Magazine 71: 272-274 [273].
Type data:
Holotype BMNH, Malanda, QLD.
Paratype(s) MMUE 2 specimens.

Taxonomic Decision for Synonymy

Hincks, W.D. 1949. Systematic and synonymic notes on Passalidae (Col.). Annals and Magazine of Natural History 12 2: 56-64 [64]

Distribution

States

Queensland

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
01-Jul-2020	SCARABAEOIDEA	23-Apr-2014	MODIFIED
		12-Feb-2010	(import)

Genus Gonatas Kaup, 1871

Gonatas Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [50].
Type species:
Passalus naviculator Percheron, 1844 by subsequent designation, see Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [108].

Distribution

States

Queensland

Extra Distribution Information

Oriental Region.

IBRA

Australian Alps (AA), Arnhem Coast (ARC), Arnhem Plateau (ARP), Avon Wheatbelt (AW), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Ben Lomond (BEL), Broken Hill Complex (BHC), Burt Plain (BRT), Central Arnhem (CA), Carnarvon (CAR), Channel Country (CHC), Central Kimberley (CK), Central Mackay Coast (CMC), Coolgardie (COO), Cobar Peneplain (CP), Central Ranges (CR), Cape York Peninsula (CYP), Daly Basin (DAB), Darwin Coastal (DAC), Desert Uplands (DEU), Dampierland (DL), Davenport Murchison Ranges (DMR), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Esperance Plains (ESP), Eyre Yorke Block (EYB), Finke (FIN), Flinders Lofty Block (FLB), Flinders (FLI), Gascoyne (GAS), Gawler (GAW), Gibson Desert (GD), Gulf Fall and Uplands (GFU), Geraldton Sandplains (GS), Great Sandy Desert (GSD), Gulf Coastal (GUC), Gulf Plains (GUP), Great Victoria Desert (GVD), Hampton (HAM), Jarrah Forest (JF), Kanmantoo (KAN), King (KIN), Little Sandy Desert (LSD), MacDonnell Ranges (MAC), Mallee (MAL), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Murchison (MUR), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), Northern Kimberley (NK), NSW North Coast (NNC), NSW South Western Slopes (NSS), Nullarbor (NUL), Ord Victoria Plain (OVP), Pine Creek (PCK), Pilbara (PIL), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Stony Plains (STP), Sturt Plateau (STU), Swan Coastal Plain (SWA), Tanami (TAN), Tasmanian Central Highlands (TCH), Tiwi Cobourg (TIW), Tasmanian Northern Midlands (TNM), Tasmanian Northern Slopes (TNS), Tasmanian South East (TSE), Tasmanian Southern Ranges (TSR), Tasmanian West (TWE), Victoria Bonaparte (VB), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Warren (WAR), Wet Tropics (WT), Yalgoo (YAL) ; Qld: Australian Alps (AA), Arnhem Coast (ARC), Arnhem Plateau (ARP), Avon Wheatbelt (AW), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Ben Lomond (BEL), Broken Hill Complex (BHC), Burt Plain (BRT), Central Arnhem (CA), Carnarvon (CAR), Channel Country (CHC), Central Kimberley (CK), Central Mackay Coast (CMC), Coolgardie (COO), Cobar Peneplain (CP), Central Ranges (CR), Cape York Peninsula (CYP), Daly Basin (DAB), Darwin Coastal (DAC), Desert Uplands (DEU), Dampierland (DL), Davenport Murchison Ranges (DMR), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Esperance Plains (ESP), Eyre Yorke Block (EYB), Finke (FIN), Flinders Lofty Block (FLB), Flinders (FLI), Gascoyne (GAS), Gawler (GAW), Gibson Desert (GD), Gulf Fall and Uplands (GFU), Geraldton Sandplains (GS), Great Sandy Desert (GSD), Gulf Coastal (GUC), Gulf Plains (GUP), Great Victoria Desert (GVD), Hampton (HAM), Jarrah Forest (JF), Kanmantoo (KAN), King (KIN), Little Sandy Desert (LSD), MacDonnell Ranges (MAC), Mallee (MAL), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Murchison (MUR), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), Northern Kimberley (NK), NSW North Coast (NNC), NSW South Western Slopes (NSS), Nullarbor (NUL), Ord Victoria Plain (OVP), Pine Creek (PCK), Pilbara (PIL), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Stony Plains (STP), Sturt Plateau (STU), Swan Coastal Plain (SWA), Tanami (TAN), Tasmanian Central Highlands (TCH), Tiwi Cobourg (TIW), Tasmanian Northern Midlands (TNM), Tasmanian Northern Slopes (TNS), Tasmanian South East (TSE), Tasmanian Southern Ranges (TSR), Tasmanian West (TWE), Victoria Bonaparte (VB), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Warren (WAR), Wet Tropics (WT), Yalgoo (YAL)

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal
Papua New Guinea
Solomon Islands

Distribution References

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [96]

General References

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [96] (for extralimital synonymy)

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Gonatas germari (Kaup, 1868)

Aceraius germari Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [30].
Type data:
Holotype unknown (as Festland von Asien).
Secondary source:
Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [51].

Distribution

States

Queensland

Extra Distribution Information

Oriental Region.

IBRA

Qld: Brigalow Belt North (BBN), Brigalow Belt South (BBS), Channel Country (CHC), Central Mackay Coast (CMC), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Gulf Plains (GUP), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Nandewar (NAN), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- Queensland
Papua New Guinea

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Gravely, F.H. 1914. An account of the Oriental Passalidae (Coleoptera) based primarily on the collection in the Indian Museum. Memoirs of the Indian Museum 3: 177-353 [301] (extralimital synonymy)

History of changes

Note that this list may be incomplete for dates prior to September 2013.

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		12-Feb-2010	(import)

Species Gonatas minimus Kuwert, 1891

Gonatas minimus Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [169].
Type data:
Syntype(s) MNHP, Ceram and Ambon (as Amboina), Indonesia.

Distribution

Extra Distribution Information

Locality in Australia unknown.

IBRA

Original AFD Distribution Data

Australian Region

Australia
Papua New Guinea
Solomon Islands

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [108] (Australian distribution record)

History of changes

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		12-Feb-2010	(import)

Species Gonatas naviculator (Percheron, 1844)

Passalus naviculator Percheron, A.R. 1844. Monographie des Passales. (Second supplément). Revue et Magasin de Zoologie (Paris) 2 6: pls 134-135 (13 pp.) [1].
Type data:
Syntype(s) Java.
Secondary source:
Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [97].

Distribution

Extra Distribution Information

Locality in Australia unknown; Oriental Region.

IBRA

Original AFD Distribution Data

Australian Region

Australia
Papua New Guinea

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103-124 [121] (extralimital synonymy)

Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349 [315] (Australian record)

History of changes

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		12-Feb-2010	(import)

Species Gonatas pumilio (Kaup, 1868)

Aceraius pumilio Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 4: 1-31 [6].
Type data:
Holotype 3 specimens, Ambon (as Amboina), Indonesia.

Distribution

States

Queensland

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal
Papua New Guinea

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Gonatas schellongi Kuwert, 1891

Gonatas schellongi Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [169].
Type data:
Holotype MNHP, New Guinea (as Neuguinea).
Gonatas albertisi Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [169].
Type data:
Syntype(s) MNHP, Cape York (as Cap York), QLD, New Britain (as Neubritannien) and New Guinea (as Neuguinea).

Taxonomic Decision for Synonymy

Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103-124 [122]

Distribution

States

Queensland

Extra Distribution Information

Oriental Region, Melanesia: New Guinea.

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Gravely, F.H. 1914. An account of the Oriental Passalidae (Coleoptera) based primarily on the collection in the Indian Museum. Memoirs of the Indian Museum 3: 177-353 [300] (taxonomy)

History of changes

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		12-Feb-2010	(import)

Genus Labienus Kaup, 1871

Labienus Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [39].
Type species:
Eriocnemis ptox Kaup, 1871 by subsequent designation, see Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [103].

Distribution

States

Queensland

Extra Distribution Information

Oriental Region.

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland
Papua New Guinea

Distribution References

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [92]

General References

History of changes

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		12-Feb-2010	(import)

Species Labienus inaequalis Gravely, 1918

Labienus inaequalis Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [104].
Type data:
Syntype(s) IM 2 specimens, Hattam and Arfak, Irian Jaya (as Dutch New Guinea), Indonesia.
Labienus trigonophorus proximus Hincks, W.D. 1933. On the species of Labienus (Col. Passalidae) related to L. trigonophorus, Zang. Stylops 2: 156-158 [158].
Type data:
Holotype MMUE, Mt Goliath, 5000–7000 ft, about 139ºE, Irian Jaya (as Central Dutch New Guinea), Indonesia.

Taxonomic Decision for Synonymy

Hincks, W.D. 1935. Notes on Passalidae (Coleoptera). No. 5 Synonymic notes, with the descriptions of a new genus and species. Entomologist's Monthly Magazine 71: 50-54 [52]

Distribution

States

Queensland

Extra Distribution Information

New Guinea.

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Genus Leptaulax Kaup, 1868

Leptaulax Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [11].
Type species:
Passalus dentatus Fabricius, 1792 by subsequent designation, see Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [112].

Distribution

Extra Distribution Information

Palaearctic and Oriental Regions.

IBRA

Original AFD Distribution Data

Australian Region

Australia
Papua New Guinea

Distribution References

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [99]

General References

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Leptaulax bicolor (Fabricius, 1801)

Passalus bicolor Fabricius, J.C. 1801. Systema Eleutheratorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Kiliae : Bibliopol. Acad. Vol. 1 xxiv 506 pp. [256].
Type data:
Syntype(s) ZMK 2 specimens, Sumatra.

Distribution

Extra Distribution Information

Locality in Australia unknown.

IBRA

Original AFD Distribution Data

Australian Region

Australia
Papua New Guinea

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Gardner, J.C.M. 1935. Immature stages of Indian Coleoptera (16) (Scarabaeoidea). Indian Forest Records Entomol. ns 1: 1-33 [5] (larval description)

History of changes

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		12-Feb-2010	(import)

Species Leptaulax dentatus (Fabricius, 1792)

Passalus dentatus Fabricius, J.C. 1792. Entomologia Systematica Emendata et Aucta. Secundum, Classes, Ordines, Genera, Species, adjectis synonimis [sic], locis, observationibus, descriptionibus. Hafniae : Christ. Gottl. Proft. Vol. 1(1) xx 330 pp. [241].
Type data:
Syntype(s) ZMK 2 specimens, Insula Guadeloupe.

Distribution

Extra Distribution Information

Locality in Australia unknown.

IBRA

Original AFD Distribution Data

Australian Region

Australia
Papua New Guinea

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Genus Mastachilus Kaup, 1868

Mastachilus Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [19].
Type species:
Passalus polyphyllus W.S. Macleay, 1826 by subsequent designation, see Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [97] (considered as the first fixation of a type).
Mastochilus Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 4: 1-31 [31] [unjustified emendation and junior objective synonym of Mastachilus Kaup, 1868].
Type species:
Passalus polyphyllus W.S. Macleay, 1826 by subsequent designation, see Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [97] (considered as the first fixation of the type).
Episphenoides Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [165].
Type species:
Passalus australasicus Percheron, 1841 by subsequent designation, see Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [87] (considered as the first fixation of a type).

Taxonomic Decision for Synonymy

Distribution

States

New South Wales, Queensland, Victoria

IBRA

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: SE coastal
- Queensland
- Victoria
Papua New Guinea

Distribution References

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [87]

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Mastachilus australasicus (Percheron, 1841)

Passalus australasicus Percheron, A.R. 1841. Révision critique et supplément à la monographie du genre Passalus. Revue et Magasin de Zoologie (Paris) 2 3: 1-48 [6].
Type data:
Syntype(s) whereabouts unknown, Australia.
Secondary source:
Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [47].
Mastachilus macleayi Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [20].
Type data:
Holotype Australia (as Neuholland).
Episphenoides mulleri Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [165].
Type data:
Holotype MNHP, Australia (as Neuholland).
Episphenoides perinvitus Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [165].
Type data:
Holotype MNHP, Australia (as Neuholland).
Episphenoides distans Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349 [328].
Type data:
Status unknown, MNHP, Australia (as Neu Holland).

Taxonomic Decision for Synonymy

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [88]

Distribution

States

New South Wales, Queensland, Victoria

IBRA

NSW, Qld, Vic: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Broken Hill Complex (BHC), Channel Country (CHC), Central Mackay Coast (CMC), Cobar Peneplain (CP), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Flinders (FLI), Gulf Plains (GUP), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), NSW North Coast (NNC), NSW South Western Slopes (NSS), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales
- Queensland
- Victoria

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Mastachilus lottinii (Boisduval, 1835)

Passalus lottinii Boisduval, J.A. 1835. Faune Entomologique de l'Océan Pacifique, avec l'illustration des insects nouveaux recueillis pendent de voyage. 2me Partie. Coléoptères et autres Ordres. In, D'Urville, D., César, J. S. (ed.). Voyage de Découvertes de l'Astrolabe exécuté par ordre du Roi, pendant les annés 1826–1829, sous le commandement de M.J. Dumont d'Urville. Paris : J. Tastu Vol. 7 716 pp. [243] [Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103–124 [117–118] did not apply priority and regarded this as a junior synonym of Episphenoides quaestionis Kuwert, 1891; on the basis of current evidence Cassis, G. & Weir, T. A. 1992. Passalidae. pp. 20–30 in Houston, W. W. K. (ed). Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Canberra : AGPS Vol. 9 [27] considered them as different species].
Type data:
Status unknown, whereabouts unknown, Australia (as Nouvelle-Hollande).
Secondary source:
Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103-124; Cassis, G. & Weir, T. A. 1992. Passalidae. pp. 20-30 in Houston, W.W.K. (ed.). Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Canberra : AGPS Vol. 9 544 pp.

Distribution

States

New South Wales

IBRA

NSW: NSW North Coast (NNC), Sydney Basin (SB), South East Corner (SEC), South Eastern Queensland (SEQ)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: SE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103-124 [117] (alternative taxonomic arrangement)

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Mastachilus parvifrons (Kuwert, 1891)

Episphenoides parvifrons Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [165].
Type data:
Holotype MNHP, Australia (as Neuholland).

Distribution

Extra Distribution Information

Locality in Australia unknown.

IBRA

Original AFD Distribution Data

Australian Region

Australia

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Mastachilus polyphyllus (W.S. Macleay, 1826)

Passalus polyphyllus Macleay, W.S. 1826. Annulosa. Catalogue of insects collected by Captain King, R.N. pp. 438-457 in King, P.P. (ed.). Narrative of a Survey of the Intertropical and Western Coasts of Australia. Performed between the years 1818 and 1822; with Appendix B. London : John Murray Vol. 2 viii 637 pp., 9 pls. [Date published 15 Apr 1826: imprint 1827] [publication date 15 April 1826 Common, I.F.B. & Moulds, M.S. 1973. The date of publication of Captain King's narrative of a survey of the intertropical and western coasts of Australia. Journal of the Society for the Bibliography of Natural History 6: 257–259 ("an initial small issue was published in the first half of 1826 and a second much larger issue in 1827")] [439].
Type data:
Status unknown, ANIC(MMUS), Australia.
Passalus sexdentatus Eschscholtz, J.F. 1829. Dissertatio de coleopterorum genere Passalus. Mémoires de la Société Impériale des Naturalistes de Moscou ns 7: 13-28 [23].
Type data:
Syntype(s) Australia.
Secondary source:
Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [88].
Passalus hexaphyllus Boisduval, J.A. 1835. Faune Entomologique de l'Océan Pacifique, avec l'illustration des insects nouveaux recueillis pendent de voyage. 2me Partie. Coléoptères et autres Ordres. In, D'Urville, D., César, J. S. (ed.). Voyage de Découvertes de l'Astrolabe exécuté par ordre du Roi, pendant les annés 1826–1829, sous le commandement de M.J. Dumont d'Urville. Paris : J. Tastu Vol. 7 716 pp. [241].
Type data:
Status unknown, Australia (as Nouvelle-Hollande).
Passalus impressicollis Boheman, C.H. 1858. Kongliga Svenska Fregatten Eugenies Resa omkring Jorden unter Befäll af C.A. Virgin 1851–53. Vetenskapliga Jakttagelser etc. (Coleoptera). Stockholm : Norstedt Vol. 1 112 pp. [40].
Type data:
Holotype NHRM, Sydney, NSW (as Sidney, Nova Hollandia).
Mastochilus abdominipunctatus Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349 [335].
Type data:
Holotype ZMB, Sydney (as Port Jackson), NSW.

Taxonomic Decision for Synonymy

Distribution

States

New South Wales, Queensland

IBRA

NSW, Qld: Brigalow Belt North (BBN), Brigalow Belt South (BBS), Channel Country (CHC), Central Mackay Coast (CMC), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Gulf Plains (GUP), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Nandewar (NAN), NSW North Coast (NNC), Sydney Basin (SB), South East Corner (SEC), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: SE coastal
- Queensland

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Mastachilus quaestionis (Kuwert, 1891)

Episphenoides quaestionis Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [165].
Type data:
Status unknown, MNHP, Australia (as Neuholland).
Mastochilus capitalis Blackburn, T. 1900. On some new genera and species of Australian Coleoptera. Proceedings of the Royal Society of Victoria 12: 206-233 [Date published Apr. 30, 1900] [209].
Type data:
Holotype BMNH, Richmond River district, NSW.

Taxonomic Decision for Synonymy

Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103-124 [117]

Distribution

States

New South Wales

IBRA

NSW: NSW North Coast (NNC), Sydney Basin (SB), South East Corner (SEC), South Eastern Queensland (SEQ)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales: SE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Genus Pharochilus Kaup, 1868

Pharochilus Kaup, J.J. 1868. Prodromus zu einer Monographie der Passaliden. Coleopterologische Hefte 3: 4-32 [20].
Type species:
Passalus dilatatus Dalman, 1817 by subsequent designation, see Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [8] (considered as the first fixation of a type).

Distribution

States

New South Wales, Queensland, Tasmania, Victoria

IBRA

Australian Alps (AA), Arnhem Coast (ARC), Arnhem Plateau (ARP), Avon Wheatbelt (AW), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Ben Lomond (BEL), Broken Hill Complex (BHC), Burt Plain (BRT), Central Arnhem (CA), Carnarvon (CAR), Channel Country (CHC), Central Kimberley (CK), Central Mackay Coast (CMC), Coolgardie (COO), Cobar Peneplain (CP), Central Ranges (CR), Cape York Peninsula (CYP), Daly Basin (DAB), Darwin Coastal (DAC), Desert Uplands (DEU), Dampierland (DL), Davenport Murchison Ranges (DMR), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Esperance Plains (ESP), Eyre Yorke Block (EYB), Finke (FIN), Flinders Lofty Block (FLB), Flinders (FLI), Gascoyne (GAS), Gawler (GAW), Gibson Desert (GD), Gulf Fall and Uplands (GFU), Geraldton Sandplains (GS), Great Sandy Desert (GSD), Gulf Coastal (GUC), Gulf Plains (GUP), Great Victoria Desert (GVD), Hampton (HAM), Jarrah Forest (JF), Kanmantoo (KAN), King (KIN), Little Sandy Desert (LSD), MacDonnell Ranges (MAC), Mallee (MAL), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Murchison (MUR), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), Northern Kimberley (NK), NSW North Coast (NNC), NSW South Western Slopes (NSS), Nullarbor (NUL), Ord Victoria Plain (OVP), Pine Creek (PCK), Pilbara (PIL), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Stony Plains (STP), Sturt Plateau (STU), Swan Coastal Plain (SWA), Tanami (TAN), Tasmanian Central Highlands (TCH), Tiwi Cobourg (TIW), Tasmanian Northern Midlands (TNM), Tasmanian Northern Slopes (TNS), Tasmanian South East (TSE), Tasmanian Southern Ranges (TSR), Tasmanian West (TWE), Victoria Bonaparte (VB), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Warren (WAR), Wet Tropics (WT), Yalgoo (YAL) ; NSW, Qld, Tas, Vic: Australian Alps (AA), Arnhem Coast (ARC), Arnhem Plateau (ARP), Avon Wheatbelt (AW), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Ben Lomond (BEL), Broken Hill Complex (BHC), Burt Plain (BRT), Central Arnhem (CA), Carnarvon (CAR), Channel Country (CHC), Central Kimberley (CK), Central Mackay Coast (CMC), Coolgardie (COO), Cobar Peneplain (CP), Central Ranges (CR), Cape York Peninsula (CYP), Daly Basin (DAB), Darwin Coastal (DAC), Desert Uplands (DEU), Dampierland (DL), Davenport Murchison Ranges (DMR), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Esperance Plains (ESP), Eyre Yorke Block (EYB), Finke (FIN), Flinders Lofty Block (FLB), Flinders (FLI), Gascoyne (GAS), Gawler (GAW), Gibson Desert (GD), Gulf Fall and Uplands (GFU), Geraldton Sandplains (GS), Great Sandy Desert (GSD), Gulf Coastal (GUC), Gulf Plains (GUP), Great Victoria Desert (GVD), Hampton (HAM), Jarrah Forest (JF), Kanmantoo (KAN), King (KIN), Little Sandy Desert (LSD), MacDonnell Ranges (MAC), Mallee (MAL), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Murchison (MUR), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), Northern Kimberley (NK), NSW North Coast (NNC), NSW South Western Slopes (NSS), Nullarbor (NUL), Ord Victoria Plain (OVP), Pine Creek (PCK), Pilbara (PIL), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Stony Plains (STP), Sturt Plateau (STU), Swan Coastal Plain (SWA), Tanami (TAN), Tasmanian Central Highlands (TCH), Tiwi Cobourg (TIW), Tasmanian Northern Midlands (TNM), Tasmanian Northern Slopes (TNS), Tasmanian South East (TSE), Tasmanian Southern Ranges (TSR), Tasmanian West (TWE), Victoria Bonaparte (VB), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Warren (WAR), Wet Tropics (WT), Yalgoo (YAL)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales
- Queensland: NE coastal
- Tasmania
- Victoria

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Pharochilus dilatatus (Dalman, 1817)

Passalus dilatatus Dalman, J.W. 1817. Sistens descriptiones novarum specierum. pp. 1-144 in Schönherr, C.J. (ed.). Appendix ad C. J. Schönherr Synonymiam Insectorum. Scaris : Lewerentziana Vol. I(3) 266 pp. [144].
Type data:
Status unknown, RMNH, Australia (as Nova Hollandia).
Passalus crenistrius Boisduval, J.A. 1835. Faune Entomologique de l'Océan Pacifique, avec l'illustration des insects nouveaux recueillis pendent de voyage. 2me Partie. Coléoptères et autres Ordres. In, D'Urville, D., César, J. S. (ed.). Voyage de Découvertes de l'Astrolabe exécuté par ordre du Roi, pendant les annés 1826–1829, sous le commandement de M.J. Dumont d'Urville. Paris : J. Tastu Vol. 7 716 pp. [244].
Type data:
Status unknown, Australia (as Nouvelle-Hollande).
Passalus chevrolati Percheron, A.R. 1835. Monographie des Passales et des genres qui en ont été séparés; accompagnée de planches dessinées par l'auteur, ou toutes les espèces ont été figurées. Paris : Mercklein iii 107 pp. 7 pls. [26].
Type data:
Syntype(s) whereabouts unknown, Australia.
Secondary source:
Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [89].
Passalus cancrus Percheron, A.R. 1844. Monographie des Passales. (Second supplément). Revue et Magasin de Zoologie (Paris) 2 6: pls 134-135 (13 pp.) [4].
Type data:
Syntype(s) whereabouts unknown, Australia.
Secondary source:
Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [89].
Tiberius caffer Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [164] [Gravely, F.H. 1914. An account of the Oriental Passalidae (Coleoptera) based primarily on the collection in the Indian Museum. Mem. Indian Mus. 3: 177–353 (194, 197) considers the type locality erroneous and that the species is undoubtedly an Australian taxon].
Type data:
Holotype MNHP, South Africa (as Caffraria).
Secondary source:
Gravely, F.H. 1914. An account of the Oriental Passalidae (Coleoptera) based primarily on the collection in the Indian Museum. Memoirs of the Indian Museum 3: 177-353.

Taxonomic Decision for Synonymy

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [89]

Distribution

States

New South Wales, Queensland

IBRA

NSW, Qld: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Broken Hill Complex (BHC), Channel Country (CHC), Central Mackay Coast (CMC), Cobar Peneplain (CP), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Gulf Plains (GUP), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Nandewar (NAN), New England Tablelands (NET), NSW North Coast (NNC), NSW South Western Slopes (NSS), Riverina (RIV), Sydney Basin (SB), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales
- Queensland

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Pharochilus nitidulus (Macleay, 1871)

Mastochilus nitidulus Macleay, W.J. 1871. Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales 2: 79-205 [174].
Type data:
Holotype AM, Gayndah, QLD.

Distribution

States

Queensland

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Pharochilus puncticollis (Macleay, 1871)

Mastochilus puncticollis Macleay, W.J. 1871. Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales 2: 79-205 [175].
Type data:
Holotype AM, Gayndah, QLD.

Distribution

States

Queensland

IBRA

Original AFD Distribution Data

Australian Region

Australia
- Queensland: NE coastal

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Pharochilus punctiger (Gravely, 1918)

Mastochilus (Pharochilus) punctiger Gravely, F.H. 1918. A contribution towards the revision of the Passalidae of the world. Memoirs of the Indian Museum 7: 1-144 [99].
Type data:
Syntype(s) IM 2 specimens, NSW.

Distribution

States

New South Wales

IBRA

NSW: Australian Alps (AA), Brigalow Belt South (BBS), Broken Hill Complex (BHC), Channel Country (CHC), Cobar Peneplain (CP), Darling Riverine Plains (DRP), Murray Darling Depression (MDD), Mulga Lands (ML), Nandewar (NAN), New England Tablelands (NET), NSW North Coast (NNC), NSW South Western Slopes (NSS), Riverina (RIV), Sydney Basin (SB), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Pharochilus rugiceps (Hope & Westwood, 1845)

Passalus dilatatus Percheron, A.R. 1835. Monographie des Passales et des genres qui en ont été séparés; accompagnée de planches dessinées par l'auteur, ou toutes les espèces ont été figurées. Paris : Mercklein iii 107 pp. 7 pls. [29] [junior homonym of Passalus dilatatus Dalman, 1817].
Type data:
Syntype(s) whereabouts unknown, ?Australia.
Secondary source:
Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [90].
Passalus rugiceps Hope, F.W. & Westwood, J.O. 1845. A Catalogue of the Lucanoid Coleoptera in the collection of the Rev. F.W. Hope, M.A., F.R.S., etc., together with Descriptions of the new species therein contained. London : J.C. Bridgewater 31 pp. [28].
Type data:
Status unknown, OUM, Port Phillip, VIC (as New Holland).
Passalus politus Burmeister, H.C.C. 1847. Handbuch der Entomologie. Coleoptera Lamellicornia Xylophila et Pectinicornia. Berlin : T.C.F. Enslin Vol. 5 viii 584 pp. [465].
Type data:
Holotype MLUH, TAS (as Van-Diemens Land).
Pharochilus politus languidilabris Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [166].
Type data:
Holotype MNHP, Australia (as Neuholland).
Pharochilus bilineatopunctatus Kuwert, A.F. 1891. Systematische Übersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift 1891: 161-192 [166].
Type data:
Holotype MNHP, Australia (as Neuholland).
Pharochilus brevidentatus Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349 [332].
Type data:
Holotype MNHP, Australia (as Neu Holland).
Pharochilus compar Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349 [333].
Type data:
Holotype MNHP, Australia (as Neu Holland).

Taxonomic Decision for Synonymy

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [89]

Distribution

States

New South Wales, Queensland, Tasmania, Victoria

IBRA

NSW, Qld, Tas, Vic: Australian Alps (AA), Brigalow Belt North (BBN), Brigalow Belt South (BBS), Ben Lomond (BEL), Broken Hill Complex (BHC), Channel Country (CHC), Central Mackay Coast (CMC), Cobar Peneplain (CP), Cape York Peninsula (CYP), Desert Uplands (DEU), Darling Riverine Plains (DRP), Einasleigh Uplands (EIU), Flinders (FLI), Gulf Plains (GUP), King (KIN), Murray Darling Depression (MDD), Mitchell Grass Downs (MGD), Mount Isa Inlier (MII), Mulga Lands (ML), Nandewar (NAN), Naracoorte Coastal Plain (NCP), New England Tablelands (NET), NSW North Coast (NNC), NSW South Western Slopes (NSS), Riverina (RIV), Sydney Basin (SB), South East Coastal Plain (SCP), South East Corner (SEC), South Eastern Highlands (SEH), South Eastern Queensland (SEQ), Simpson Strzelecki Dunefields (SSD), Tasmanian Central Highlands (TCH), Tasmanian Northern Midlands (TNM), Tasmanian Northern Slopes (TNS), Tasmanian South East (TSE), Tasmanian Southern Ranges (TSR), Tasmanian West (TWE), Victorian Midlands (VM), Victorian Volcanic Plain (VVP), Wet Tropics (WT)

Original AFD Distribution Data

Australian Region

Australia
- New South Wales
- Queensland
- Tasmania
- Victoria

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

General References

Schuster, J.C. & Reyes-Castillo, P. 1981. New World genera of Passalidae (Coloptera): a revision of larvae. Anales de la Escuela Nacional de Ciencias Biologicas, Mexico 25: 79-116 (taxonomy)

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Pharochilus scutellonotus Kuwert, 1898

Pharochilus scutellonotus Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349 [333].
Type data:
Syntype(s) MNHP 3 specimens, Australia (as Neu Holland).
Pharochilus pararius Kuwert, A.F. 1898. Die Passaliden dichotomisch bearbeitet. Novitates Zoologicae 5: 259-349 [334].
Type data:
Holotype MNHP, Australia (as Neu Holland).

Taxonomic Decision for Synonymy

Dibb, J.R. 1938. Synopsis of Australian Passalidae (Coleoptera). Transactions of the Royal Entomological Society of London 87: 103-124 [119]

Distribution

Extra Distribution Information

Locality in Australia unknown.

IBRA

Original AFD Distribution Data

Australian Region

Australia

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Genus Protomocoelus Zang, 1905

Protomocoelus Zang, R. 1905. Bemerkenswerte Passaliden aus der Sammlung des Zoologischen Instituts in Strassburg. Deutsche Entomologische Zeitschrift 1905: 105-112 [154] [nom. nov. for Pelops Kaup, 1871].
Pelops Kaup, J.J. 1871. Monographie der Passaliden. Berliner Entomologische Zeitschrift 15: 1-125 [37] [junior homonym of Pelops Gistl, 1834 (Coleoptera)].
Type species:
Passalus australis Boisduval, 1835 by monotypy.

Distribution

Extra Distribution Information

Oriental Region.

IBRA

Original AFD Distribution Data

Australian Region

Australia
New Caledonia
Papua New Guinea
Solomon Islands

Distribution References

Hincks, W.D. & Dibb, J.R. 1935. Passalidae. pp. 1-118 in Schenkling, S. (ed.). Coleopterorum Catalogus. Lucanidae — Passalidae — Scarabaeidae I. Berlin : W. Junk Vol. XIX Pt 142. [95]

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Species Protomocoelus australis (Boisduval, 1835)

Passalus australis Boisduval, J.A. 1835. Faune Entomologique de l'Océan Pacifique, avec l'illustration des insects nouveaux recueillis pendent de voyage. 2me Partie. Coléoptères et autres Ordres. In, D'Urville, D., César, J. S. (ed.). Voyage de Découvertes de l'Astrolabe exécuté par ordre du Roi, pendant les annés 1826–1829, sous le commandement de M.J. Dumont d'Urville. Paris : J. Tastu Vol. 7 716 pp. [246].
Type data:
Status unknown, Dori, Indonesia (as Doreï, Nouvelle-Guinée).

Distribution

Extra Distribution Information

Solomon Islands, New Guinea, New Caledonia; locality in Australia unknown.

IBRA

Original AFD Distribution Data

Australian Region

Australia
New Caledonia
Solomon Islands

Ecological Descriptors

Adult: terrestrial, volant.

Extra Ecological Information

Larvae and adults inhabit and feed on rotten wood.

History of changes

Note that this list may be incomplete for dates prior to September 2013.

Published	As part of group	Action Date	Action Type	Compiler(s)
		12-Feb-2010	(import)

Family PASSALIDAE

Compiler and date details

Introduction

Diagnosis

Diagnosis References

General References

History of changes

Subfamily Aulacocyclinae

History of changes

Genus Aulacocyclus Kaup, 1868

Type species:

Type species:

Taxonomic Decision for Synonymy

Distribution

States

Extra Distribution Information

IBRA

Original AFD Distribution Data

Australian Region

Distribution References

History of changes

Species Aulacocyclus aliicornis Kuwert, 1897

Type data:

Type data:

Taxonomic Decision for Synonymy

Distribution

States

IBRA

Original AFD Distribution Data

Australian Region

Ecological Descriptors

Extra Ecological Information

History of changes

Species Aulacocyclus collaris Blackburn, 1896

Type data:

Distribution

States

IBRA

Original AFD Distribution Data

Australian Region

Ecological Descriptors

Extra Ecological Information

History of changes

Species Aulacocyclus deyrolli Kaup, 1868

Type data:

Distribution

States

Extra Distribution Information

IBRA

Original AFD Distribution Data

Australian Region

Ecological Descriptors

Extra Ecological Information

History of changes

Species Aulacocyclus edentulus (W.S. Macleay, 1826)

Type data:

Type data:

Type data:

Secondary source:

Type data:

Taxonomic Decision for Synonymy

Distribution

States

IBRA

Original AFD Distribution Data

Australian Region

Ecological Descriptors

Extra Ecological Information

General References

History of changes

Species Aulacocyclus fracticornis Kuwert, 1891

Type data:

Type locality references:

Secondary source:

Type data:

Secondary source:

Type data:

Taxonomic Decision for Synonymy

Distribution

States